1995-2000 Ph.D. in Biology, Universitat Autònoma de Barcelona (Spain). 2001-2003 EMBO Postdoctoral Fellow, The Salk Institute for Biological Studies (USA). 2003-2006 Postdoctoral Fellow, Emory University (USA). 2006-2009 Ramon y Cajal Researcher, Centre for Genomic regulation (Spain) 2010-pres. ICREA Research Professor, Universitat Autònoma de Barcelona (Spain).